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Stem

Eccentric

False chanterelle (Hygrophoropsis aurantiaca)

Mycological descriptions often use the term eccentric stem (eccentric stipe). This stem position is in between perfectly central (as in porcini) and strictly lateral (as in the oyster mushroom).

If you turn such a mushroom cap-side down, you'll see that the stem is attached neither in the middle of the cap nor at its very edge. It is offset (shifted) from the center to one side.

6 species in the database.

Why does the stem shift?

As David Arora explains, an eccentric stem is most often the mushroom's response to the spatial conditions it grows in.

A mushroom always tries to hold its cap perfectly horizontal (so the spores can fall freely downward). If it grows on a steep slope, on a leaning tree trunk or at an angle out of a crack in a log, its stem will bend and attach to the cap off-center, compensating for the tilt of the substrate.

Typical species with an eccentric stem

Although eccentricity is often an individual response, for some mushrooms it is the norm for the species:

1. Sawgills (Neolentinus): Large mushrooms that grow on dead wood (especially pine). Their very dense, tough stem is almost always attached to the cap eccentrically. Their gills have finely serrated (saw-toothed) edges, which is their main feature.

2. Oyster mushroom (Pleurotus ostreatus): Although we usually call the oyster mushroom's stem lateral, in young mushrooms or those growing on horizontal surfaces it is often precisely eccentric (the cap grows unevenly around the stem).

3. Dryad's saddle (Cerioporus squamosus): Its large, scaly cap always rests on a thick eccentric (or lateral) stem that blackens at the base.

Questions and answers

If I find a porcini with an eccentric stem, is it a mutant?
No, it's not a mutation. If a porcini (which normally has a perfectly central stem) grew on a very steep ravine bank or had to grow around a thick root underground, its stem may bend and shift away from the center of the cap. It's simply an ecological adaptation that has no effect whatsoever on its edibility or safety.
Are there eccentric amanitas?
Extremely rarely. Amanitas grow on the ground, usually on level patches of forest litter. Their architecture (cap, ring, volva) calls for a symmetrical, central structure for maximum stability. Eccentricity in amanitas is possible only as a mechanical deformation during growth (for example, if the cap pushed up against a root).

Sources

  1. 1.Зерова М.Я. та ін. Визначник грибів України. 1979
  2. 2.Roger Phillips. Mushrooms (Roger Phillips)
  3. 3.David Arora. Mushrooms Demystified
  4. 4.Index Fungorum