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Stem

Absent

Absent — mushrooms with the trait «Absent»

For many macrofungi, especially those that specialize in breaking down dead or living wood, the stem turned out to be an entirely unnecessary organ. Mycologists call such fungi sessile or bracket fungi.

When a mushroom has no stem, its cap is attached to the substrate (the tree) directly by its side or back.

24 species in the database.

Why did they give up the stem?

As M.Ya. Zerova notes, the main purpose of a stem is to lift the mushroom above the ground so the wind can carry its spores. But if a fungus is already growing on a tree trunk two meters up, there's no point spending resources on building a stem — it's high enough as it is. The cap simply grows onto the bark, forming a sturdy horizontal shelf whose gills or tubes face straight down.

The main stemless fungi

The largest group of sessile fungi is the polypores (Polyporaceae). Their fruit bodies are often shaped like a horse's hoof, a half-circle or a burl.

1. Tinder fungus (Fomes fomentarius): The classic gray "hoof" on the trunks of old birches. It has no stem at all. It is fixed rock-solid to the bark and is very hard and woody.

2. Splitgill (Schizophyllum commune): A very small, widespread fungus that grows on dead branches. It looks like tiny hairy gray shells with no stems. Interestingly, its gills can split in two in dry weather.

3. Beefsteak fungus (Fistulina hepatica): This unique edible fungus grows on oaks and resembles a piece of raw liver in shape and color. It has no stem; it attaches to the trunk by its narrowed side (sometimes with a very short pseudo-root).

Balls and stars

Besides wood-dwelling fungi, the gasteromycetes that grow on the ground also have no stems:

- Puffballs (Lycoperdon): Some species (the stump puffball, for example) have a small sterile base (a pseudostem), but no true stem — they're simply balls sitting on the substrate.

- Earthstars (Geastrum): They are ball-shaped, and their outer skin splits open into a star that stays lying on the ground without any stem.

Questions and answers

If the oyster mushroom grows on wood, why does it have a stem (a lateral one), while a bracket fungus doesn't?
It comes down to the fungus's evolutionary path and texture. The oyster mushroom is a soft, gilled mushroom that grows fast and lives for a few days. Polypores (many of them) are perennial fungi. Their caps have the texture of wood and grow in layers year after year. Forming a stem for such a heavy, perennial, woody structure would be inefficient, so a bracket fungus "grows onto" the tree with its entire broad base for maximum strength.

Sources

  1. 1.Зерова М.Я. та ін. Визначник грибів України. 1979
  2. 2.Roger Phillips. Mushrooms (Roger Phillips)
  3. 3.David Arora. Mushrooms Demystified
  4. 4.Index Fungorum